000150372 001__ 150372 000150372 005__ 20251017144629.0 000150372 0247_ $$2doi$$a10.1093/zoolinnean/zlab113 000150372 0248_ $$2sideral$$a128678 000150372 037__ $$aART-2022-128678 000150372 041__ $$aeng 000150372 100__ $$aRotatori, Filippo Maria 000150372 245__ $$aReappraisal and new material of the holotype of Draconyx loureiroi (Ornithischia: Iguanodontia) provide insights on the tempo and modo of evolution of thumb-spiked dinosaurs 000150372 260__ $$c2022 000150372 5060_ $$aAccess copy available to the general public$$fUnrestricted 000150372 5203_ $$aThe Upper Jurassic Lourinha Formation is well known for its rich assemblage of fossil vertebrates. In this formation, ornithopod dinosaurs are represented by two iguanodontian species, Eousdryosaurus nanohallucis and Draconyx loureiroi. We recently became aware of unreported material belonging to the holotype of Draconyx loureiroi, consisting of partially articulated manual elements. We here re-describe the holotype specimen ML 357, including the newly discovered material. The specimen was subjected to CT-scanning and its surface data used to assess anatomical characters. Linear measurements of metatarsal III were used to estimate the body length of the specimen. The Draconyx loureiroi holotype was included in two datasets and analysed with maximum parsimony and Bayesian inference approaches to estimate evolutionary rates among Iguanodontia. We present evidence that Draconyx loureiroi is a valid taxon nested in Styracosterna and is clearly diagnosable by a unique combination of characters. Both maximum parsimony and Bayesian inference indicate high evolutionary rates across the Jurassic/Cretaceous transition for the base of Iguanodontia. Length estimation suggests that Draconyx loureiroi was a relatively small, bipedal and possibly cursorial animal. Given its basal phyletic position, we interpret this bauplan was the ancestral condition for Styracosterna, that only later in the Cretaceous evolved into giant quadrupedal forms. 000150372 540__ $$9info:eu-repo/semantics/openAccess$$aby-nc-nd$$uhttps://creativecommons.org/licenses/by-nc-nd/4.0/deed.es 000150372 590__ $$a2.8$$b2022 000150372 591__ $$aZOOLOGY$$b12 / 176 = 0.068$$c2022$$dQ1$$eT1 000150372 592__ $$a1.055$$b2022 000150372 593__ $$aEcology, Evolution, Behavior and Systematics$$c2022$$dQ1 000150372 593__ $$aAnimal Science and Zoology$$c2022$$dQ1 000150372 594__ $$a6.4$$b2022 000150372 655_4 $$ainfo:eu-repo/semantics/article$$vinfo:eu-repo/semantics/acceptedVersion 000150372 700__ $$0(orcid)0000-0002-7210-1033$$aMoreno-Azanza, Miguel$$uUniversidad de Zaragoza 000150372 700__ $$aMateus, Octavio 000150372 7102_ $$12000$$2655$$aUniversidad de Zaragoza$$bDpto. Ciencias de la Tierra$$cÁrea Paleontología 000150372 773__ $$g20 (2022), 1-32$$pZool. j. Linn. Soc.$$tZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY$$x0024-4082 000150372 8564_ $$s6873433$$uhttps://zaguan.unizar.es/record/150372/files/texto_completo.pdf$$yPostprint 000150372 8564_ $$s1563687$$uhttps://zaguan.unizar.es/record/150372/files/texto_completo.jpg?subformat=icon$$xicon$$yPostprint 000150372 909CO $$ooai:zaguan.unizar.es:150372$$particulos$$pdriver 000150372 951__ $$a2025-10-17-14:25:43 000150372 980__ $$aARTICLE