Mycorrhizal competition release and microbial dynamics innative and non-native Tuber melanosporum habitats Benucci Gian Maria Niccolò Garcia-Barreda Sergi Sanchez Sergio Marco Pedro Universidad de Zaragoza De Miguel Ana Maria Le Tacon Francois Marozzi Giorgio Baciarelli Failini Leonardo Eslick Harry Elliott Todd F. Deveau Aurelie Murat Claude Donnini Domizia Bonito Gregory 2026 research-article