doi:10.1038/s41598-026-49942-wengAbián, DavidBernad, JorgeIlarri, SergioTrillo-Lado, RaquelIndividual and collective gains from cooperation and reciprocity in a dynamic-network Prisoner’s Dilemma driven by extraversion, openness, and agreeablenessART-2026-149247How do stable personality differences shape cooperation when social ties can form and dissolve? We model a repeated Prisoner’s Dilemma on an endogenous network in which three continuous Big Five traits map to transparent local mechanisms: Extraversion sets a target number of partners, Openness determines how broadly agents search beyond friends-of-friends, and Agreeableness sets a baseline willingness to cooperate. At each encounter, agents combine this baseline with the partner’s directly observed history; there are no trait labels, gossip, or global reputations. Ties form when agents are under-connected and are cut when they become over-connected, with cuts prioritising partners who have defected more often. We vary network size (N=30–200), population composition, and the balance between trait-driven and history-driven behaviour. Three robust patterns emerge. First, cooperate first, then reciprocate—high initial willingness to cooperate combined with history-sensitive response—produces systems that are simultaneously more prosperous, fairer, and safer. Second, personality has predictable conditional effects: Agreeableness helps when history matters but hurts when behaviour is mostly trait-driven; Extraversion amplifies the environment; Openness has little net payoff effect. Third, the network reorganises accordingly: degree assortativity stays near zero, whereas agreeable agents increasingly connect to one another when cooperation takes hold.2026http://zaguan.unizar.es/record/17125210.1038/s41598-026-49942-whttp://zaguan.unizar.es/record/171252oai:zaguan.unizar.es:171252info:eu-repo/grantAgreement/ES/AEI/PID2020-113037RB-I00info:eu-repo/grantAgreement/ES/AEI/PID2020-113903RB-I00info:eu-repo/grantAgreement/ES/DGA/T42-23Rinfo:eu-repo/grantAgreement/ES/DGA/T64-23RScientific reports (Nature Publishing Group) (2026), [14 pp.]byhttps://creativecommons.org/licenses/by/4.0/deed.esinfo:eu-repo/semantics/openAccess