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<articles>
<article xmlns:xlink="http://www.w3.org/1999/xlink/">
  <front>
    <journal-meta>
      <journal-title>NATURE COMMUNICATIONS</journal-title>
      <abbrev-journal-title>NATURE COMMUNICATIONS</abbrev-journal-title>
      <issn/>
    </journal-meta>
    <article-meta>
      <title-group>
        <article-title>Integrative epigenome-wide analysis demonstrates that DNA methylation may mediate genetic risk in inflammatory bowel disease</article-title>
      </title-group>
      <contrib-group>
        <contrib contrib-type="author">
          <name>
            <surname>Ventham</surname>
            <given-names>N.T.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Kennedy</surname>
            <given-names>N.A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Adams</surname>
            <given-names>A.T.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Kalla</surname>
            <given-names>R.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Heath</surname>
            <given-names>S.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>O''Leary</surname>
            <given-names>K.R.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Drummond</surname>
            <given-names>H.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Wilson</surname>
            <given-names>D.C.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Gut</surname>
            <given-names>I.G.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Nimmo</surname>
            <given-names>E.R.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Satsangi</surname>
            <given-names>J.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Lauc</surname>
            <given-names>G.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Campbell</surname>
            <given-names>H.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>McGovern</surname>
            <given-names>D.P.B.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Annese</surname>
            <given-names>V.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Zoldoš</surname>
            <given-names>V.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Permberton</surname>
            <given-names>I.K.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Wuhrer</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Kolarich</surname>
            <given-names>D.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Fernandes</surname>
            <given-names>D.L.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Theorodorou</surname>
            <given-names>E.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Merrick</surname>
            <given-names>V.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Spencer</surname>
            <given-names>D.I.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Gardner</surname>
            <given-names>R.A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Doran</surname>
            <given-names>R.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Shubhakar</surname>
            <given-names>A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Boyapati</surname>
            <given-names>R.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Rudan</surname>
            <given-names>I.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Lionetti</surname>
            <given-names>P.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Akmacic</surname>
            <given-names>I.T.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Krištic</surname>
            <given-names>J.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Vuckovic</surname>
            <given-names>F.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Štambuk</surname>
            <given-names>J.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Novokmet</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Pucic-Bakovic</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Gornik</surname>
            <given-names>O.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Andriulli</surname>
            <given-names>A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Cantoro</surname>
            <given-names>L.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Sturniolo</surname>
            <given-names>G.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Fiorino</surname>
            <given-names>G.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Manetti</surname>
            <given-names>N.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Latiano</surname>
            <given-names>A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Kohn</surname>
            <given-names>A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>D''Incà</surname>
            <given-names>R.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Danese</surname>
            <given-names>S.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Arnott</surname>
            <given-names>I.D.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Noble</surname>
            <given-names>C.L.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Lees</surname>
            <given-names>C.W.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Shand</surname>
            <given-names>A.G.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Ho</surname>
            <given-names>G.T</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Dunlop</surname>
            <given-names>M.G.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Murphy</surname>
            <given-names>L.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Gibson</surname>
            <given-names>J.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Evenden</surname>
            <given-names>L.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Wrobel</surname>
            <given-names>N.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Gilchrist</surname>
            <given-names>T.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Fawkes</surname>
            <given-names>A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Kammeijer</surname>
            <given-names>G.S.M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Clerc</surname>
            <given-names>F.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>De Haan</surname>
            <given-names>N.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Vojta</surname>
            <given-names>A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Samaržija</surname>
            <given-names>I.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Markulin</surname>
            <given-names>D.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Klasic</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Dobrinic</surname>
            <given-names>P.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Aulchenko</surname>
            <given-names>Y.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Van</surname>
            <given-names>Den Heuve</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Jonkers</surname>
            <given-names>D.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Pierik</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Vatn</surname>
            <given-names>S.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Ricanek</surname>
            <given-names>P.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Jahnsen</surname>
            <given-names>J.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>You</surname>
            <given-names>P.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Sølvernes</surname>
            <given-names>J.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Frengen</surname>
            <given-names>A.B.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Tannæs</surname>
            <given-names>T.M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Moen</surname>
            <given-names>A.E.F.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Dahl</surname>
            <given-names>F.A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Lindstrøm</surname>
            <given-names>J.C.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Ekeland</surname>
            <given-names>G.S.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Detlie</surname>
            <given-names>T.E.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Keita</surname>
            <given-names>A.V.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Söderholm</surname>
            <given-names>J.D.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Hjortswang</surname>
            <given-names>H.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Halfvarson</surname>
            <given-names>J.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Bergemalm</surname>
            <given-names>D.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Gomollón</surname>
            <given-names>F.</given-names>
          </name>
          <aff>
            <institution>Universidad de Zaragoza</institution>
          </aff>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>D'Amato</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Törkvist</surname>
            <given-names>L.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Hjelm</surname>
            <given-names>F.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Gullberg</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Nordberg</surname>
            <given-names>N.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Ocklind</surname>
            <given-names>A.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Pettersson</surname>
            <given-names>E.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Ekman</surname>
            <given-names>D.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Sundell</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Modig</surname>
            <given-names>E.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Veillard</surname>
            <given-names>A.C.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Schoemans</surname>
            <given-names>R.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Poncelet</surname>
            <given-names>D.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Sabatel</surname>
            <given-names>C.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Gut</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Bayes</surname>
            <given-names>M.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Casén</surname>
            <given-names>C.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Lindahl</surname>
            <given-names>T.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Ciemniejewska</surname>
            <given-names>E.</given-names>
          </name>
        </contrib>
        <contrib contrib-type="author">
          <name>
            <surname>Vatn</surname>
            <given-names>M.H.</given-names>
          </name>
        </contrib>
      </contrib-group>
      <pub-date pub-type="pub">
        <year>2016</year>
      </pub-date>
      <volume/>
      <fpage/>
      <lpage/>
      <self-uri xlink:href="http://zaguan.unizar.es/record/58481"/>
      <self-uri xlink:href="http://zaguan.unizar.es/record/58481/files/texto_completo.pdf"/>
      <self-uri xlink:href="http://zaguan.unizar.es/record/58481/files/texto_completo.jpg?subformat=icon"/>
    </article-meta>
    <abstract/>
  </front>
  <article-type>research-article</article-type>
  <ref/>
</article>

</articles>