Prion Prion Scrapie strain typing of brain and lymph node-derived isolates in ovinized models reveals mixture of substrains with distinct pathobiological properties Barrio T Universidad de Zaragoza Filali H Universidad de Zaragoza Otero A Sheleby-Elias J Marin B Universidad de Zaragoza Vidal E Torres J.M Groschup M Andreoletti O Badiola J.J Bolea R. 2019 research-article